2005 CIC bioGUNE, Bilbao, Spain.
2006-2010 Ramón y Cajal Fellow.
2002-2005 Marie Curie Postdoctoral Fellow at Cancer Research UK, London, UK.
1997-2002 EMBO, Human Frontiers, NATO Postdoctoral Fellow at EMBL, Monterotondo, Italy.
1992-1996 PhD in Biology at Harvard University, Massachusetts, USA.
1990-1992 MA in Biology at Harvard University, Massachusetts, USA.
1988-1990 BA in Biology at Oberlin College, Ohio, USA.